NICHE DIFFERENTIATION IN TROPICAL FOREST

Equilibrium hypotheses for the maintenance of tropical diversity generally invoke some form of niche differentiation. This hypothesis is based upon the idea that ecologically similar species are unable to coexist unless they have developed different patterns of habitat distribution and/or resource use. Within this framework, the more specialized the resource requirements of each species are, the more species can be packed into a given habitat. Niche differences among tropical animals are generally related to the type of food resources utilized, or spatial or temporal differences in habitat use.

For example, otherwise ecologically similar animals can differ in terms of height of activity in the canopy, or the time of day they are active. In contrast, all plant species utilize essentially the same set of basic resources, namely: light, water, carbon dioxide, physical space, and nutrients such as nitrogen, phosphorus and potassium.

However, plant species can differ in terms of more subtle ecological characteristics, such as the efficiency of resource use, tolerance of physiological stress, or dependence on specific pollinators, seed-dispersers or root symbionts. Many studies of tropical forest trees have emphasized differences in the "regeneration niche", or the resources and conditions required by seedlings and saplings to successfully establish in the forest. An important distinction is made between ‘pioneer’ tree species that can grow rapidly in large canopy openings or cleared areas, and "late-successional" or "primary forest" tree species that can establish under low light conditions in the understorey. Other kinds of niche differences among tropical trees include ‘structural niche’ differences related to the size reached by adult trees, and differences in ‘habitat preference’ related to soil characteristics and hydrology.

CLIMATE AND BIOLOGICAL PRODUCTIVITY OF THE TROPICS

The warm, wet, and relatively aseasonal climate of the tropics is apparently more favourable for maintaining higher diversity than anywhere else in the world. But why is this the case? One simple idea is that the high solar energy inputs and productivity of tropical regions result in greater numbers of species that can be supported energetically.


However, it is not entirely clear why a small number, or even one species, could not monopolize most or all of the incoming solar energy. Another idea is that climate stability is the main factor promoting species diversification and coexistence. In the harsher temperate and polar regions, species must be able to tolerate drastic fluctuations in seasonal temperatures. Species occurring in habitats nearer the poles are therefore adapted to a wider range of local environments in order to survive the winter months.

As a consequence, one expects a narrower range of adaptation to environmental conditions and narrower latitudinal and altitudinal distributions in the tropics, a hypothesis sometimes referred to as ‘Rapoport’s rule’. The more limited ranges of species in the tropics may allow for greater ‘species packing’ compared to temperate or boreal regions.

Although this idea has received much research attention, recent analyses give only equivocal support, at best, for Rapoport’s rule. On the other hand, it is clear that the relatively aseasonal nature of tropical rain forests allows for the evolution of highly varied and complex species interactions. This complexity itself contributes to the overall species diversity found in the tropics. For example, "dependent" ecological forms, such as specialist herbivores or predators, only persist in the community if their host species is present.

HISTORICAL IMPORTANCE OF TROPICAL FORESTS IN BIOLOGY

Tropical forests have played a central role in the conceptual development of biology from the time of the major biological expeditions that began at the close of the eighteenth century.

Alexander Von Humbolt initiated the study of plant ecology on his voyages through South America in the early nineteenth century. While climbing Mount Chimborazo in the Andes, Von Humbolt characterized the vegetation changes with climate as he ascended. These early observations on plant distributions provided the foundation of the field of biogeography. The most significant development in biological thinking inspired by tropical forests was the theory of evolution by natural selection. Charles Darwin and Alfred Russel Wallace independently derived this theory as a result of their scientific voyages in the tropics during the mid nineteenth century. Darwin’s inspiration was his exploration of various parts of South America as the naturalist aboard the Beagle beginning in 1831.

Wallace conducted expeditions in both South America (1848–1852) and the Malay Archipelago (1854–1862), where he characterized two sets of fauna distinct to the different parts of the archipelago, a division now known as Wallace’s Line. For both Darwin and Wallace, the high diversity of species in the tropics, and particularly patterns of diversification associated with island groups, allowed an appreciation of evolutionary relationships not apparent in the poorer faunas of the temperate zone. The high diversity of the tropics continues to be an important inspiration and testing ground for ideas in biology, particularly in the fields of ecology and evolutionary biology.

DISTRIBUTION PATTERNS TROPICAL RAINFOREST

The Tropical forests exist with some essential facts of geography and climatic systems. By definition, ‘the tropics’ lie between 23o N and 23o S latitude, the area within which the sun lies directly overhead at some point in its seasonal progression.

The flux of solar energy within this region is high, due to the fact that incoming sunlight is projected at a 90o angle to the surface of the earth. This high solar energy flux results in a high rate of water evaporation over the tropical oceans and of evapotranspiration over tropical land surfaces.



The result is a rising column of warm, moist air at tropical latitudes. As this air rises it cools as a result of the gradient in air pressure through the atmosphere (adiabatic cooling). Water condenses out from this air mass, generating rainfall. The air mass ultimately dries out, and is carried poleward from the tropics as a part of wind circulation patterns known as Hadley cells.

At subtropical latitudes near 23–30o N and S, the now-dry air mass descends, creating a region of high pressure that corresponds closely to the world distribution of deserts. As a consequence of these climatic circulation patterns, the earth’s equatorial zone is warm and wet, corresponding to the broad band of tropical forests found along the earth’s equatorial axis.

WHAT IS THE TROPICS | Tropics Are The Regions Of The Earth

Tropics are the regions of the earth that lie within about 1,600 miles (2,570 kilometers) north and 1,600 miles south of the equator. Two imaginary lines, the Tropic of Cancer and the Tropic of Capricorn, form the boundaries of the tropics. The Tropic of Cancer is 23° 27' north of the equator, and the Tropic of Capricorn is 23° 27' south of the equator. These lines mark the northernmost and southernmost places on the earth where the sun ever shines directly overhead.

Most places in the tropics have warm to hot temperatures the year around. Tropical places near sea level are hot because every day the sun's rays shine almost straight down at noon. Such direct rays produce higher temperatures than do slanted rays.

The temperature does not change much in the tropics because the amount of daylight differs little from season to season. At the equator, the sun shines about 12 hours a day. At the edges of the tropics, daylight varies from about 101/2 hours a day in winter to about 131/2 hours a day in summer. Places at the edges of the tropics have cool periods in winter. Tropical places that are located at high altitudes are cool because the temperature drops about 31/2 °F per 1,000 feet (2 °C per 300 meters) of elevation.

Many tropical areas have definite rainy and dry seasons. Most places near the equator get much rain during all seasons and are covered by tropical rain forests. Farther to the north and south, one or two short dry seasons occur yearly. Such areas have forests of trees that lose their leaves during these dry seasons. Areas even farther from the equator have one long dry season each year. These areas are covered by savannas (grasslands with scattered trees and shrubs).